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Although genes were known to exist on chromosomes, chromosomes are composed of both protein and DNA, and scientists did not know which of the two is responsible for inheritance.In 1928, Frederick Griffith discovered the phenomenon of transformation (see Griffith's experiment): dead bacteria could transfer genetic material to "transform" other still-living bacteria.A classic example is two seeds of genetically identical corn, one placed in a temperate climate and one in an arid climate.While the average height of the two corn stalks may be genetically determined to be equal, the one in the arid climate only grows to half the height of the one in the temperate climate due to lack of water and nutrients in its environment.James Watson and Francis Crick determined the structure of DNA in 1953, using the X-ray crystallography work of Rosalind Franklin and Maurice Wilkins that indicated DNA has a helical structure (i.e., shaped like a corkscrew).

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The intracellular or extracellular environment of a cell or organism may switch gene transcription on or off.Genetics has given rise to a number of subfields, including epigenetics and population genetics.Organisms studied within the broad field span the domain of life, including bacteria, plants, animals, and humans.Although the structure of DNA showed how inheritance works, it was still not known how DNA influences the behavior of cells.In the following years, scientists tried to understand how DNA controls the process of protein production.

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